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Oolon Colluphid
01 Apr 2009, 11:49 AM
PNAS vol. 106 no. 13 pp. 5235-5240 (31 March 2009 )

Retroposon analysis and recent geological data suggest near-simultaneous divergence of the three superorders of mammals

Nishihara, Maruyama and Okada

As a consequence of recent developments in molecular phylogenomics, all extant [currently alive] orders of placental mammals have been grouped into 3 lineages: Afrotheria [noted for their puffball hair], Xenarthra, and Boreotheria, which originated in Africa, South America, and Laurasia, respectively.

Despite this advancement, the order of divergence of these 3 lineages remains unresolved. Here, we performed extensive retroposon analysis with mammalian genomic data.

Surprisingly, we identified a similar number of informative retroposon loci that support each of 3 possible phylogenetic hypotheses: the basal position [diverged first] for Afrotheria (22 loci), Xenarthra (25 loci), and Boreotheria (21 loci).

This result indicates that the divergence of the placental common ancestor into the 3 lineages occurred nearly simultaneously.

Thus, we examined whether these molecular data could be integrated into the geological context by incorporating recent geological data. We obtained firm evidence that complete separation of Gondwana into Africa and South America occurred 120 ± 10 Ma. Accordingly, the previous reported time frame (division of Pangea into Gondwana and Laurasia at 148–138 Ma and division of Gondwana at 105 Ma) cannot be used to validate mammalian divergence order. Instead, we use our retroposon results and the recent geological data to propose that near-simultaneous divisions of continents leading to isolated Africa, South America, and Laurasia caused nearly concomitant divergence of the ancient placental ancestor into 3 lineages, Afrotheria, Xenarthra, and Boreotheria, ≈120 Ma.
Open Access article (http://www.pnas.org/content/106/13/5235.full) (whole thing for free)

Steviepinhead
01 Apr 2009, 09:13 PM
<([Bracketed comments in the abstract are Oolon's, I presume {puffball hair, indeed! :cool: }.])>

Notta
02 Apr 2009, 01:05 AM
I like how the article uses the term 'simultaneously' to cover a period of a couple of million years......

espritch
02 Apr 2009, 02:35 AM
I like how the article uses the term 'simultaneously' to cover a period of a couple of million years......

For a geologist, anything less than 10 million years is an eye blink.

lpetrich
02 Apr 2009, 10:50 AM
<([Bracketed comments in the abstract are Oolon's, I presume {puffball hair, indeed! :cool: }.])>
Making them all look like poodles. *joke*

The relationships between mammalian orders have long been controversial, and it's nice to see those relationships finally being resolved.

And what's interesting here is that various specializations evolved separately in the various groups; the ancestors of each group had been primitive and unspecialized.

An old taxon, Ungulata, putting elephants among Perissodactyla (odd-toed ungulates) and Artiodactyla (even-toed ungulates), is "polyphyletic", and thus not a legitimate taxon. Elephants are in Afrotheria and the other ungulates in Boreotheria.

Specializing for eating ants and termites evolved four times:
Boreotheria: Pholidota: pangolin
Xenarthra: South American anteater
Afrotheria: Tubulidentata: aardvark
Monotremata: spiny anteater

Going aquatic has evolved three times:
Boreotheria: Carnivora: Pinnipedia: seals and the like
Boreotheria: Artiodactyla: Cetacea: dolphins and whales
Afrotheria: Sirenia: manatees and the like

Burrowing and living underground have also evolved several times.

A rhinoceros-like build and head shape has evolved several times:
Boreotheria: Perissodactyla: rhinoceros
Boreotheria: Perissodactyla: titanotheres
Boreotheria: Artiodactyla: hippopotamus (hornless)
Boreotheria(?): Uintatherium
Afrotheria: Arsinoitherium
Marupialia: Diprotodon
Dinosauria: ceraptopsians

Elephants have a similar build but short heads, pigs and tapirs have similar body and head forms but are much smaller, etc.

Etc.

Some terminology:

Location | Taxon | Other Two
Africa | Afrotheria | Exafroplacentalia
South America | Xenarthra | Epitheria
Laurasia | Boreotheria | Atlantogenata
Laurasia = North America + Eurasia

DMB
02 Apr 2009, 02:06 PM
<channelling creationist> It all happened very fast, as I have already said. The animals got out of the ark and were whizzed away on the floating continents at high speed. </channelling creationist>

Robert
13 Apr 2010, 05:02 PM
If these mammals were spread out across Gondwana before the split, then when the split occurred they would already be in the areas of Gondwana which we now called South America, Africa and Europe.
So when the split occurred they would already be in those areas. So it would not matter if the split occurred simultaneously or not.